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Conceptualization, W.H.H and R.A.; Methodology, W.H.H. and R.A.; Investigation, W.H.H., J.G., M.H., A.W., R.M.V., B.T.K., and S.J.I.; Resources, P.L., J-X.L., W.J.L., and G.J.F.; Writing - Original Draft, W.H.H. and R.A.; Writing - Review and Editing, W.H.H., J.G., A.W., S.J.I., H.T.K., and R.A.; Visualization, W.H.H.; Supervision, R.A. and H.T.K.; Funding Acquisition, W.H.H. and R.A.
The authors thank Alice Kamphorst, Koichi Araki, Timothy Hoang, and Robert Karaffa and Kametha Fife of the Emory University School of Medicine Flow Cytometry Core.
Rafi Ahmed holds patents on the PD-1 pathway.
W.H.H. is a Cancer Research Institute Irvington Fellow supported by the Cancer Research Institute. H.T.K. is supported by grant K99CA197804 from the National Cancer Institute of the National Institutes of Health (NIH). R.A. is supported by grants R01AI030048 and P01AI056299 from the National Institute of Allergy and Infectious Diseases of the NIH. The Yerkes Nonhuman Primate Genomics Core is supported in part by ORIP/OD P51OD011132. The content is solely the responsibility of the authors and does not necessarily represent the official views of the National Institutes of Health.
2019