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To whom correspondence should be addressed. Email:kressler@mclean.harvard.edu Edited by Joseph E. LeDoux, New York University, New York, NY, and approved September 8, 2015 (received for review March 12, 2015)
F.G.M., B.G.D., and K.J.R. designed research; F.G.M. and B.G.D. performed research; F.G.M., B.G.D., and K.J.R. contributed new reagents/analytic tools; F.G.M., B.G.D., and K.J.R. analyzed data; and F.G.M., B.G.D., and K.J.R. wrote the paper.
We thank the animal care staff in the Yerkes Neuroscience Vivarium for assistance with animal husbandry.
We also thank Kenneth McCullough, Stephanie Maddox, and Sunayana Banerjee for assistance with ensuring double-blindness of data acquisition and analysis; Kevin Ding for assistance with olfactory fear behavior; and members of the K.J.R. laboratory for thoughtful and helpful feedback on the manuscript and support.
The authors declare no conflict of interest.
This article is a PNAS Direct Submission.
This work was supported by the following sources of funding: the National Institutes of Mental Health (1R01MH096764) (to K.J.R.), the Howard Hughes Medical Institute, and Ruth L. Kirstein National Research Service Award (NRSA) Predoctoral Fellowship F31 MH105237-01 (to F.G.M.).
This project was also funded by the Office of Research Infrastructure Programs/OD P51OD011132 [formerly National Center for Research Resources (NCRR) P51RR000165].